Hila May Lab
The Biohistory & Evolutionary Medicine Laboratory


Publications
Y Zaidner, M Prévost, R Shahack-Gross, L Weissbrod, R Yeshurun, N Porat, G Guérin, N Mercier, A Galy, C Pécheyran, G Barbotin, C Tribolo, H Valladas, D White, R Timms, S Blockley, A Frumkin, D Gaitero-Santos, S Ilani, S Ben-Haim, A Pedergnana, AV Pietraszek, P García, C Nicosia, S Lagle, O Varoner, C Zeigen, D Langgut, O Crouvi, S Borgel, R Sarig, H May, I Hershkovitz. Evidence from Tinshemet Cave in Israel suggests behavioural uniformity across Homo groups in the Levantine mid-Middle Palaeolithic circa 130,000–80,000 years ago. Nature Human Behaviour, 11:1-6, 2025. https://doi.org/10.1038/s41562-025-02110-y
S Peleg, R Pelleg-Kallevag, G Dar, N Steinberg, Z Lenzner, H May. The effect of Scheuermann's kyphosis on rib cage morphology: A skeletal study. Annals of Anatomy-Anatomischer Anzeiger, 1;257:152348, 2025.
A Hadad, Y Bar-Ziv, H May. New insights on femoral head trabecular bone directionality in elderly humans using a micro-CT-based analysis. Scientific Reports 15, 24228, 2025. https://doi.org/10.1038/s41598-025-05991-1
E Kedar, D Ezra, R Pelleg-Kallevag, D Stein, N Peled, H May, I Hershkovitz. Capturing the cervical spine shape: Angular measurements versus geometric morphometric methods. Clinical Anatomy, 38(3), 228–238, 2025. https://doi.org/10.1002/ca.24166
Y Smorgick, R Pelleg-Kallevag, D Lindner, Y Anekstein, S Goldstein, H May. Vertebral body density role in determining vertebral osteoporotic fracture type and its progression. Clinical Anatomy, 38(1), 97–104, 2025. https://doi.org/10.1002/ca.24219
D Megreli, A Pokhojaev, H May, DZ Bar, R Sarig. Inter-and intra-period dietary variations revealed by dental microwear texture analysis in Holocene Levantine populations. Scientific Reports, 2025. https://doi.org/10.1038/s41598-025-30282-0
R Yeshurun, G Hartman, H May, F Rivals, KM Crater Gershtein, C Zeigen, Y Zaidner. Archaic humans in the Middle Palaeolithic Levant conducted planned and selective intercepts of aurochs, but not mass hunting. Scientific Reports, 26;15(1):42237, 2025. https://doi.org/10.1038/s41598-025-26274-9
A Eirikh-Rose, K Zutovski, D Ackerfeld, Y Asscher, H May, Y Makoviychuk, L Namdar, Y Gordover, L Sapir-Hen, HC Schechter, H Ashkenazi. Nahal Yarmuth 38: A PPNB and PN Site in Central Israel. Palestine Exploration Quarterly, 21:1-36, 2025. https://doi.org/10.1080/00310328.2025.2562700
A Pokhojaev, HL Levine, T Sella-Tunis, R Sarig, H May. Author Correction: Changes in human mandibular shape during the Terminal Pleistocene-Holocene Levant. Sci Rep. 2025 Aug 15;15(1):29933. doi: 10.1038/s41598-025-15216-0. Erratum for: Sci Rep. 2019 Jun 19;9(1):8799. doi: 10.1038/s41598-019-45279-9. PMID: 40817398; PMCID: PMC12356901.
HL Levine, N Shvalb, A Pokhojaev, S Francis, R Pelleg-Kallevag, V Roul, JJ Hublin, F Rühli, H May. Author Correction: Evolutionary roots of the risk of hip fracture in humans. Commun Biol. 2025 Jul 18;8(1):1068. doi: 10.1038/s42003-025-08498-7. Erratum for: Commun Biol. 2023 Mar 17;6(1):283. doi: 10.1038/s42003-023-04633-4. PMID: 40681791; PMCID: PMC12274277.
JM López-Rey, I Crevecoeur, H May, D Nadel, CA Palancar, M Gómez-Recio, D García-Martínez, M Bastir. Fossil ribcages of Homo sapiens provide new insights into modern human evolution. Communications Biology, 10;8(1):1038, 2025. https://doi.org/10.1038/s42003-025-08472-3
M Ullman, H May, S Ya'aran, B Langford, I Hershkovitz, L Chavoinik, N Marom, DL Gnezdilov, A Frumkin, U Davidovich. Death and Depths: Exploring Early Fifth Millennium bce Ritual Performance in Har Sifsof Cave, Upper Galilee (Israel). Cambridge Archaeological Journal, 35(2):380-99, 2025. doi:10.1017/S0959774325000022
R Pelleg-Kallevag, S Borgel, E Kedar, N Peled, H May. Changes in the shape of the lumbar curve during growth. Bone Joint Res, 14(1):58-68, 2025. doi:10.1302/2046-3758.141.BJR-2024-0081.R1
S Borgel, S Francis, U Avner, G Stiebel, H May. Long live the sick nobleman! New insights into the Nabataean society from the Israeli desert. International Journal of Paleopathology, 46: 12-12, 2024.
S Bejdova, R Rmoutilova, H May, I Crevecoeur, H Rougier, J Veleminska, P Veleminsky, J Bruzek. Morphology of the early Upper Paleolithic Zlaty kun mandible, Czech Republic. American Journal of Biological Anthropology, 183: 15-16, 2024.
Q Cosnefroy, G Berillon, T Chevalier, AM Kubicka, C De Becdelievre, H May, F Marchal. Alternative biomechanical strategies of Neandertals and Homo sapiens femora during the Pleistocene. In93rd Annual Meeting of the American Association of Biological Anthropologists, 2024.
N Sala, A Pantoja-Pérez, L Martín-Francés, H May, I Hershkovitz. Life and death in the Middle Paleolithic of Qafzeh, Israel through comprehensive taphonomic study of hominin remains, 2024.
Q Cosnefroy, G Berillon, AM Kubicka, C de Becdelièvre, T Chevalier, H May, F Marchal. La courbure et les propriétés biomécaniques de la diaphyse fémorale d’Homo neanderthalensis et d’Homo sapiens révèlent-elles des stratégies bio-mécaniques distinctes pour la bipédie?. Bulletins et mémoires de la Société d’Anthropologie de Paris. BMSAP. 1;36(36 (S)), 2024.
O Barzilai, O Marder, J Tejero, A Ayalon, M Bar-Matthews, T Abulafia, R Lavi, M Goder-Goldberger, M Shemer, L Edeltin, A Wiegmann, A Frumkin, A Karasik, G Yasur, R Yeshurun, I Zohar, F Berna, M Hans, JS Goldberg, Y McDermott, L Spurlock, A Pokhojaev, W Habashi, H May, R Sarig, I Hershkovitz. Early human collective practices and symbolism in the Early Upper Paleolithic of Southwest Asia. Proc. Natl. Acad. Sci. U.S.A. 121 (51) e2404632121, 2024. DOI: https://doi.org/10.1073/pnas.2404632121.
Y Smorgick, R Pelleg‐Kallevag, D Lindner, Y Anekstein, S Goldstein, H May. Vertebral body density role in determining vertebral osteoporotic fracture type and its progression. Clinical Anatomy, 2024.
L Namdar, L Bartosiewicz, H May* , L Sapir‐Hen* . Animals' paleopathology: Implications on human–animal interaction during the intensification of farming in the Southern Levant. International Journal of Osteoarchaeology, e3333, 2024.
E Kedar, D Ezra, R Pelleg‐Kallevag, D Stein, N Peled, H May, and I Hershkovitz. Capturing the cervical spine shape: Angular measurements versus geometric morphometric methods. Clinical Anatomy, 1-11, 2024.
A Dann, A Pokhojaev, M Anton, G Yalovitsky, R Kallevag-Pelleg, and H May. Changes in human calcaneal morphology throughout the Pleistocene-Holocene Levant. Frontiers in Earth Science, 12: 1294350, 2024.
A Pantoja-Pérez, L Martín-Francés, H May, I Hershkovitz, and N Sala. New potential case of Paleolithic interpersonal violence: First taphonomic approach from Qafzeh-25, Israel. 2023.
S Francis, Y Makoviychuck, L Chavoinik, S Borgel, A Pokhojaev, V Roul, N Peled, and H May. A new method for sex estimation based on femoral cross-sectional geometry measurements and its validation using recent and ancient populations. International Journal of Legal Medicine, 6:1-3, 2023 .
A Oikonomou, Y Asscher, H May, Y Gorin-Rosen, and T Rehren. Metal prills in primary glass: A puzzling aspect of the production process. Journal of Archaeological Science: Reports, 49:103977, 2023.
HL Avni, N Shvalb, A Pokhojaev, S Francis, R Pelleg-Kallevag, V Roul, JJ Hublin, F Rühli, and H May. Evolutionary roots of the risk of hip fracture in humans. Communication Biology, 6(1): 283, 2023.
I Nudel, A Bader-Farage, N Shpack, H May, R Sarig. Age Estimation Of Archeological Populations. Journal of Archeological Dcience Using Secondary Dentin Analysis. Journal of Archaeological Science, 151: 105724, 2023.
O Bender, D Tzabag, T Gavish, N Meyrom, N Zamir, H May, R Sarig, and DZ Bar. The hidden secrets of the dental calculus: Calibration of a mass spectrometry protocol for dental calculus protein analysis. International journal of Molecular Sciences, 23(22): 14387, 2022.
A Balzau, L Albessard-Ball, AM Kubicka, A Filippo, A Beaudet, E Santos, T Bienvenu, JL Arsuage, A Bartsiokas, L Berger, JM Bermudez de Castro, M Brunet, KJ Carlson, J Daura, VG Gorgoulis, FE Grine, K Harvati, J Hawks, A Herries, JJ Hublin, J Hui, R Ives, JA Joordens, Y Kaifu, M Kouloukoussa, B Leger, D Lorfkipanidze, A Margvelashvili, J Martin, M Martinon-Torres, H May, A Mounier, A du Plessis, T Rae, C Roding, M Sanz, P Semal, D Stratford, C Striner, M Tawane, H Temming, E Tsoulkalka, J Zilhao, B Zippfel, and LT Buck. Frontal sinuses and human evolution. Science advances, 8(42): eabp9767, 2022.
SA Spyrou, G Noblesb, A Hadjikoumis, A Evin, A Hulme-Beaman, C Çakirlar, C Ameen, N Loucas, E Nikita, P Hanot, N M de Boer, A Avgousti, I Zohar, H May, Th Rehren. Digital Zooarchaeology: State of the art, challenges, prospects and synergies. Journal of Archaeological Science: Reports, 45: 103588, 2022.
O Barzilai, T Abulafia, M Shemer, H May, M Orbach, A Frumkin, R Yeshurun, R Sarig, N Porat, and I Hershkovitz. Rediscovering Geula Cave: A Middle Paleolithic cave site in northern Mt. Carmel, Israel. Quaternary International, 624: 181-197, 2022.
H May, R Sarig, A Pokhojaev, C Fornai, M Martinón-Torres, J María B de Castro, GW Weber, Y Zaidner, and I Hershkovitz. "Response to Comment on “A Middle Pleistocene Homo from Nesher Ramla, Israel”." Science 374, no. 6572: eabl5789, 2021.
S Borgel, B Latimer, Y McDermott, R Sarig, A Pokhojaev, T Abulafia, M Goder-Goldberger, O Barzilai, H May. Early Upper Paleolithic human foot bones from Manot Cave, Israel. J Hum Evol.,160: 102668, 2021.
I Hershkovitz*, H May*, R Sarig*, A Pokhojaev, D Grimaud-Hervé, E Bruner, C Fornai, R Quam, JL Arsuaga, VA Krenn, M Martinón-Torres,JM Bermúdez de Castro,L Martín-Francés, V Slon, L Albessard-Ball, A Vialet, T Schüler, G Manzi, A Profico, F Di Vincenzo, GW Weber, Y Zaidner. A Middle Pleistocene Homo from Nesher Ramla, Israel. Science, 372(6549): 1424-1428, 2021.
Y Zaidner, L Centi, M Prévost, N Mercier, C Falguères, G Guérin, H Valladas, M Richard, A Galy, C Pécheyran, O Tombret, E Pons-Branchu, N Porat, R Shahack-Gross, DE Friesem, R Yeshurun, Z Turgeman-Yaffe, A Frumkin, G Herzlinger, R Ekshtain, M Shemer, O Varoner, R Sarig*, H May*, I Hershkovitz*. Middle Pleistocene Homo behavior and culture at 140,000 to 120,000 years ago and interactions with Homo sapiens. Science, 372(6549): 1429-1433, 2021.
R Sorrentino, N.B Stephens, D Marchi, L.J.D DeMars, C Figus, E Bortolini, F Badino, J.P.P Saers, M Bettuzzi, F Boschin, G Capecchi, F Feletti, T Guarnieri, H May, M.P Morigi, W Parr, S Ricci, A Ronchitelli, J.T Stock, K.J Carlson, T.M Ryan, M.G Belcastro, S Benazzi. Unique foot posture in Neanderthals reflects their body mass and high mechanical stress. Journal of Human Evolution, 131: 103093, 2021.
W Habashi, A Bader-Farraj, N Shpack, I Beitlitum, H May, R Sarig. Three-Dimensional Surface Texture Analysis of Fluoride’s Effect on Enamel Erosion. Journal of Clinical Medicine, 10 (19): 4528, 2021.
B Medlej, H Cohen, G Dar, S Peleg, J Abbas, Y Masharawi, I Hershkovitz, H May. Defects of the femoral head‐neck junction: A new method of classification and observed frequency in Hamann‐Todd skeletal collection. International Journal of Osteoarchaeology, 31(5): 801-808, 2021.
Y Leshno Afriat, Y Edelman-Furstenberg, R Rabinovich, JA Todd, H May. Taxonomic identification using virtual palaeontology and geometric morphometerics: A case study of Jurassic nerineoidean gastropods. Paleontology, 64(2): 249-261, 2021.
D Grimaud-Hervé, L Albessard-Ball, A Pokhojaev, A Balzeau, R Sarig, B Latimer, Y McDermott, H May, I Hershkovitz. The Endocast of the Late Middle Paleolithic Manot 1 Specimen, Western Galilee, Israel. Journal of Human Evolution,160: 102734, 2021.
T Sella Tunis, H May, R Sarig, AD Vardimon, I Hershkovitz, N Spack. Are chin and symphysis morphology facial type dependent? A CT-based study. American Journal of Orthodontics and Dentofacial Orthopedics, 160(1): 84-93, 2021.
I Nudel, A Pokhojaev, BS Hausman, Y Bitterman, N Shpack, H May, R Sarig. Age estimation of fragmented human dental remains by secondary dentin virtual analysis. International Journal of Legal Medicine, 134(5): 1853-1860, 2020.
T Sella Tunis, I Hershkovitz, H May, AD Vardimon, R Sarig, Nir Shpack. Variation in Chin and Mandibular Symphysis Size and Shape in Males and Females: A CT-Based Study. International Journal of Environmental Research and Public Health, 17(12): 4249, 2020. doi: 10.3390/ijerph17124249.
K Floreanova, E Gilat, I Koren, H May. Ear infection prevalence in prehistoric and historic populations of the southern Levant: A new diagnostic method. International Journal of Osteoarchaeology, 30(4): 449-457, 2020. https://doi.org/10.1002/oa.2873).
S Peleg, R Pelleg-Kallevag, G Dar, N Steinberg, Y Masharawi, H May. New methods for sex and ancestry estimation using sternum and rib morphology. International Journal of Legal Medicine. Accepted, 134: 1519-1530, 2020. DOI: https://doi.org/10.1007/s00414-020-02266-4
BM Rothschild, D Tanke, F Rühli, A Pokhojaev, H May. Suggested Case of Langerhans Cell Histiocytosis in a Cretaceous Dinosaur. Sci. Rep., 10: 2203, 2020. DOI: https://doi.org/10.1038/s41598-020-59192-z
GW Weber, I Hershkovitz, P Gunz, S Neubauer, A Ayalone, B Latimer, M Bar-Matthews, G Yasur, O Barzilai, H May. Before the massive modern human dispersal into Eurasia: A 55,000-year-old partial cranium from Manot Cave, Israel. Quaternary International, 551: 29-39, 2020.
DOI: https://doi.org/10.1016/j.quaint.2019.10.009
A Gopher, A Eirikh-Rose, H Ashkenazi, E Marco, H May, Y Makoviychuk, L Sapir-Hen, S Galmor, HC Schechter, D Ackerfeld, G Haklay, K Zutovski. Nahal Yarmuth 38: A new and unique PPNB site in central Israel. Antiquity, 93(371): e29, 2019. DOI: https://doi.org/10.15184/aqy.2019.162
R Sarig, C Fornai, A Pokhojaev, H May, M Hans, B Latimer, O Barzilai, R Quam, GW Weber. The dental remains from the Early Upper Paleolithic of Manot Cave, Israel. J Hum Evol., 102648, 2019.
A Pokhojaev, H Avni, T Sella-Tunis, R Sarig, H May. Changes in human mandibular shape during the Terminal Pleistocene-Holocene Levant. Sci Rep., 9(1): 8799, 2019.
M Feldman, E Fernández-Domínguez, L Reynolds, D Baird, J Pearson, I Hershkovitz, H May, N Goring-Morris, M Benz, J Gresky, RA Bianco, A Fairbairn, G Mustafaoğlu, PW Stockhammer, C Posth, W Haak, C Jeong, J Krause. Late Pleistocene human genome suggests a local origin for the first farmers of central Anatolia. Nat Commun., 10(1): 1218, 2019.
H May, T Sella-Tunis, A Pokhojaev, N Peled, R Sarig. Changes in mandible characteristics during the terminal Pleistocene to Holocene Levant and their association with dietary habits. Journal of Archaeological Science: Reports, 22: 413-419, 2018.
M Edry, E Arie, H May, A Yasur-Landau. The Iron Age IIA Tombs of Area E, Tel Achziv: Between Local Traditions and the Consolidation of the Tyrian Polity. Israel Exploration Journal, 68(2): 150-181, 2018.
A Pokhojaev, W Habashi, H May, E Schulz-Kornas, N Shvalb, R Sarig. Examination of the Interproximal Wear Mechanism: Facet Morphology and Surface Texture Analysis. Journal of Dental Research, 97(13): 1145-1451, 2018. https://doi.org/10.1177/0022034518785140.
É Harney, H May, D Shalem, N Rohland, S Mallick, I Lazaridis, R Sarig, K Stewardson, S Nordenfelt, N Patterson, I Hershkovitz, D Reich. Ancient DNA from Chalcolithic Israel reveals the role of population mixture in cultural transformation. Nat Commun., 9(1): 3336, 2018.
T Sella-Tunis, A Pokhojaev, R Sarig, P O'Higgins, H May. Human mandibular shape is associated with masticatory muscle force. Sci Rep., 8(1): 6042, 2018.
I Hershkovitz, GW Weber, R Quam, M Duval, R Grün, L Kinsley, A Ayalon, M Bar-Matthews, H Valladas, N Mercier, JL Arsuaga, M Martinón-Torres, JM Bermúdez de Castro, C Fornai, L Martín-Francés, R Sarig, H May, VA Krenn, V Slon, L Rodríguez, R García, C Lorenzo, JM Carretero, A Frumkin, R Shahack-Gross, DE Bar-Yosef Mayer, Y Cui, X Wu, N Peled, I Groman-Yaroslavski, L Weissbrod, R Yeshurun, A Tsatskin, Y Zaidner, M Weinstein-Evron. The earliest modern humans outside Africa. Science, 359(6734): 456-459, 2018.
H Cohen, C Kugel, H May, B Medlej, D Stein, V Slon, T Brosh, I Hershkovitz. The influence of impact direction and axial loading on the bone fracture pattern. Forensic Sci Int., 277: 197-206, 2017.
T Tunis, R Sarig, H Cohen, B Medlej, N Peled, H May. Sex estimation using computed tomography of the mandible. Int J Legal Med., 131(6): 1691-1700, 2017.
H Cohen, C Kugel, H May, B Medlej, D Stein, V Slon, T Brosh, I Hershkovitz. The effect of impact tool geometry and soft material covering on long bone fracture patterns in children. Int J Legal Med., 131(4): 1011-1021, 2017.
J Abbas, V Slon, H May, N Peled, I Hershkovitz, K Hamoud. Paraspinal muscles density: a marker for degenerative lumbar spinal stenosis? BMC Musculoskelet Disord, 17(1): 1-8, 2016.
H Cohen, C Kugel, H May, B Medlej, D Stein, V Slon, I Hershkovitz, T Brosh. The impact velocity and bone fracture pattern: Forensic perspective. Forensic Sci Int., 266: 54-62, 2016.
H May, C Ruff. Physical burden and lower limb bone structure at the origin of agriculture in the levant. AJPA., 161(1): 26-36, 2016.
H Cohen, C Kugel, H May, B Medlej, D Stein, V Slon, I Hershkovitz, T Brosh. The impact velocity and bone fracture pattern: Forensic perspective. Forensic Sci Int., 266:54-62, 2016.
O Hay, G Dar, J Abbas, D Stein, H May, Y Masharawi, N Peled, I Hershkovitz. The Lumbar Lordosis in Males and Females, Revisited. PLos One, 10(8): e.0133685, 2015.
R Sarig, I Hershkovitz, N Shpack, H May, AD Vardimon. Rate and pattern of interproximal dental attrition. Eur J Oral Sci., 123(4): 276-81, 2015.
V Slon, D Stein, H Cohen, T Sella-Tunis, H May, I Hershkovitz. Vertebral hemangiomas: their demographical characteristics, location along the spine and position within the vertebral body. Eur Spine J., 24(10): 2189-95, 2015.
H Cohen, V Slon, A Barash, H May, B Medlej, I Hershkovitz. Assyrian attitude towards captive enemies: A 2700-year-old paleo-forensic study. Int J Osteoarcheol., 25(3): 265-80, 2015.
I Hershkovitz, HD Donoghue, DE Minnikin, H May, OY Lee, M Feldman, E Galili, M Spigelman, BM Rothschild, GK Bar-Gal. Tuberculosis origin: The Neolithic scenario. Tuberculosis, 95: S122-6, 2015.
I Hershkovitz, O Marder, A Ayalon, M Bar-Matthews, G Yasur, E Boaretto, V Caracuta, B Alex, A Frumkin, M Goder-Goldberger, P Gunz, RL Holloway, B Latimer, R Lavi, A Matthews, V Slon, DB Mayer, F Berna, G Bar-Oz, R Yeshurun, H May, MG Hans, GW Weber, O Barzilai. Levantine cranium from Manot Cave (Israel) foreshadows the first European modern humans. Nature, 520(7546): 216-9, 2015.
H May, F Rühli. Is the Shape of the Proximal Femur a Risk Factor for Degenaritve Processes of the Hip and Hip Fracture? A New Approach Based On Geometric Morphometrics Methods. The FASEB Journal., 29 (1 Supplement): 702-3, 2015.
I Hershkovitz, M Spigelman, R Sarig, DS Lim, IS Lee,CS Oh, H May, E Boaretto, YS KimS, SD Lee, N Peled, MJ Kim, T Toledano, GK Bar-Gal, DH Shin. A possible case of cherubism in a 17th-century Korean mummy.
PLoS One, 9(8): e102441, 2014.
R Sarig, I Hershkovitz, N Shvalb, T Sella-Tunis, H May, AD Vardimon. Proximal attrition facets: morphometric, demographic, and aging characteristics. Eur J Oral Sci., 122(4): 271-8, 2014.
V Slon, I Hershkovitz, H May. The value of cadaver CT scans in gross anatomy laboratory. Anat Sci Educ., 7(1): 80-2, 2014.