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A Dann, Pokhojaev, A., Anton, M., Yalovitsky, G., Kallevag-Pelleg, R., & May, H. Changes in human calcaneal morphology throughout the Pleistocene-Holocene Levant. Frontiers in Earth Science, 12: 1294350, 2024.

A Pantoja-Pérez, L Martín-Francés, H May, I Hershkovitz, N Sala. New potential case of Paleolithic interpersonal violence: First taphonomic approach from Qafzeh-25, Israel. 2023.

S Francis, Y Makoviychuck, L Chavoinik, S Borgel, A Pokhojaev, V Roul, N Peled, H May. A new method for sex estimation based on femoral cross-sectional geometry measurements and its validation using recent and ancient populations. International Journal of Legal Medicine, 6:1-3, 2023 .

A Oikonomou, Y Asscher, H May, Y Gorin-Rosen, T Rehren. Metal prills in primary glass: A puzzling aspect of the production process. Journal of Archaeological Science: Reports, 49:103977, 2023.

H L Avni, N Shvalb, A Pokhojaev, S Francis, R Pelleg-Kallevag, V Roul, J J Hublin, F Rühli, and H May. Evolutionary roots of the risk of hip fracture in humans. Communication Biology, 6(1): 283, 2023.

I Nudel, A Bader-Farage, N Shpack, H May, R Sarig. Age Estimation Of Archeological Populations. Journal of Archeological Dcience Using Secondary Dentin Analysis. Journal of Archaeological Science, 151: 105724, 2023.

O Bender, D Tzabag, T Gavish, N Meyrom, N Zamir, H May, R Sarig, and DZ Bar. The hidden secrets of the dental calculus: Calibration of a mass spectrometry protocol for dental calculus protein analysis. International journal of Molecular Sciences, 23(22): 14387, 2022.

A Balzau, L Albessard-Ball, AM Kubicka, A Filippo, A Beaudet, E Santos, T Bienvenu, JL Arsuage, A Bartsiokas, L Berger, JM Bermudez de Castro, M Brunet, KJ Carlson, J Daura, VG Gorgoulis, FE Grine, K Harvati, J Hawks, A Herries, JJ Hublin, J Hui, R Ives, JA Joordens, Y Kaifu, M Kouloukoussa, B Leger, D Lorfkipanidze, A Margvelashvili, J Martin, M Martinon-Torres, H May, A Mounier, A du Plessis, T Rae, C Roding, M Sanz, P Semal, D Stratford, C Striner, M Tawane, H Temming, E Tsoulkalka, J Zilhao, B Zippfel, and LT Buck. Frontal sinuses and human evolution. Science advances, 8(42): eabp9767, 2022.

SA Spyrou, G Noblesb, A Hadjikoumis, A Evin, A Hulme-Beaman, C Çakirlar, C Ameen, N Loucas, E Nikita, P Hanot, N M de Boer, A Avgousti, I Zohar, H May, Th Rehren. Digital Zooarchaeology: State of the art, challenges, prospects and synergies. Journal of Archaeological Science: Reports, 45: 103588, 2022.

O Barzilai, T Abulafia, M Shemer, H May, M Orbach, A Frumkin, R Yeshurun, R Sarig, N Porat, and I Hershkovitz. Rediscovering Geula Cave: A Middle Paleolithic cave site in northern Mt. Carmel, Israel. Quaternary International, 624: 181-197, 2022.

H May, R Sarig, A Pokhojaev, C Fornai, M Martinón-Torres, J María B de Castro, GW Weber, Y Zaidner, and I Hershkovitz. "Response to Comment on “A Middle Pleistocene Homo from Nesher Ramla, Israel”." Science 374, no. 6572: eabl5789, 2021.

Borgel S, Latimer B, McDermott Y, Sarig R, Pokhojaev A, Abulafia T, Goder-Goldberger M, Barzilai O, May H. Early Upper Paleolithic human foot bones from Manot Cave, Israel. J Hum Evol.,160: 102668, 2021. 

I Hershkovitz*, H May*, R Sarig*, A Pokhojaev, D Grimaud-Hervé, E Bruner, C Fornai, R Quam, JL Arsuaga, VA Krenn, M Martinón-Torres,JM Bermúdez de Castro,L Martín-Francés, V Slon, L Albessard-Ball, A Vialet, T Schüler, G Manzi, A Profico, F Di Vincenzo, GW Weber, Y Zaidner. A Middle Pleistocene Homo from Nesher Ramla, Israel. Science, 372(6549): 1424-1428, 2021.

Y Zaidner, L Centi, M Prévost, N Mercier, C Falguères, G Guérin, H Valladas, M Richard, A Galy, C Pécheyran, O Tombret, E Pons-Branchu, N Porat, R Shahack-Gross, DE Friesem, R Yeshurun, Z Turgeman-Yaffe, A Frumkin, G Herzlinger, R Ekshtain, M Shemer, O Varoner, R Sarig*, H May*, I Hershkovitz*. Middle Pleistocene Homo behavior and culture at 140,000 to 120,000 years ago and interactions with Homo sapiens. Science, 372(6549): 1429-1433, 2021.

R Sorrentino, N.B Stephens, D Marchi, L.J.D DeMars, C Figus, E Bortolini, F Badino, J.P.P Saers, M Bettuzzi, F Boschin, G Capecchi, F Feletti, T Guarnieri, H May, M.P Morigi, W Parr, S Ricci, A Ronchitelli, J.T Stock, K.J Carlson, T.M Ryan, M.G Belcastro, S Benazzi. Unique foot posture in Neanderthals reflects their body mass and high mechanical stress. Journal of Human Evolution, 131: 103093, 2021.


W Habashi, A Bader-Farraj, N Shpack, I Beitlitum, H May, R Sarig. Three-Dimensional Surface Texture Analysis of Fluoride’s Effect on Enamel Erosion. Journal of Clinical Medicine, 10 (19): 4528, 2021.

B Medlej, H Cohen, G Dar, S Peleg, J Abbas, Y Masharawi, I Hershkovitz, H May. Defects of the femoral head‐neck junction: A new method of classification and observed frequency in Hamann‐Todd skeletal collection. International Journal of Osteoarchaeology, 31(5): 801-808, 2021.

Y Leshno Afriat, Y Edelman-Furstenberg, R Rabinovich, JA Todd, H May. Taxonomic identification using virtual palaeontology and geometric morphometerics: A case study of Jurassic nerineoidean gastropods. Paleontology, 64(2): 249-261, 2021.

Grimaud-Hervé D, Albessard-Ball L, Pokhojaev A, Balzeau A, Sarig R, Latimer B, McDermott Y, May H, Hershkovitz I. The Endocast of the Late Middle Paleolithic Manot 1 Specimen, Western Galilee, Israel. Journal of Human Evolution,160: 102734, 2021

T Sella Tunis, H May, R Sarig, AD Vardimon, I Hershkovitz, N Spack. Are chin and symphysis morphology facial type dependent? A CT-based study. American Journal of Orthodontics and Dentofacial Orthopedics, 160(1): 84-93, 2021

I Nudel, A Pokhojaev, BS Hausman, Y Bitterman, N Shpack, H May, R Sarig. Age estimation of fragmented human dental remains by secondary dentin virtual analysis. International Journal of Legal Medicine, 134(5): 1853-1860, 2020.

T Sella Tunis, I Hershkovitz, H May, AD Vardimon, R Sarig, Nir Shpack. Variation in Chin and Mandibular Symphysis Size and Shape in Males and Females: A CT-Based Study. International Journal of Environmental Research and Public Health, 17(12): 4249, 2020. doi: 10.3390/ijerph17124249.

K Floreanova, E Gilat, I Koren, H May. Ear infection prevalence in prehistoric and historic populations of the southern Levant: A new diagnostic method. International Journal of Osteoarchaeology, 30(4): 449-457, 2020.

Peleg S, Pelleg Kallevag R, Dar G, Steinberg N, Masharawi Y, May H. New methods for sex and ancestry estimation using sternum and rib morphology. International Journal of Legal Medicine. Accepted, 134: 1519-1530, 2020. DOI:

Rothschild BM, Tanke D, Rühli F, Pokhojaev A, May H. Suggested Case of Langerhans Cell Histiocytosis in a Cretaceous Dinosaur. Sci. Rep., 10: 2203, 2020. DOI:

Weber G W, Hershkovitz I, Gunz P, Neubauer S, Ayalone A, Latimer B, Bar-Matthews M, Yasur G, Barzilai O, May H. Before the massive modern human dispersal into Eurasia: A 55,000-year-old partial cranium from Manot Cave, Israel. Quaternary International, 551: 29-39, 2020



Gopher A, Eirikh-Rose A, Ashkenazi H, Marco E, May H, Makoviychuk Y, Sapir-Hen L, Galmor S, Schechter H C, Ackerfeld D, Haklay G, Zutovski K. Nahal Yarmuth 38: A new and unique PPNB site in central Israel. Antiquity, 93(371): e29, 2019. DOI:

Sarig R, Fornai C, Pokhojaev A, May H, Hans M, Latimer B, Barzilai O, Quam R, Weber GW.  The dental remains from the Early Upper Paleolithic of Manot Cave, Israel. J Hum Evol., 102648, 2019.


Pokhojaev A, Avni H, Sella-Tunis T, Sarig R, May H. Changes in human mandibular shape during the Terminal Pleistocene-Holocene Levant. Sci Rep., 9(1): 8799, 2019.


Feldman M, Fernández-Domínguez E, Reynolds L, Baird D, Pearson J, Hershkovitz I, May H, Goring-Morris N, Benz M, Gresky J, Bianco RA, Fairbairn A, Mustafaoğlu G, Stockhammer PW, Posth C, Haak W, Jeong C, Krause J. Late Pleistocene human genome suggests a local origin for the first farmers of central Anatolia. Nat Commun., 10(1): 1218, 2019.


H May, T Sella-Tunis, A Pokhojaev, N Peled, R Sarig. Changes in mandible characteristics during the terminal Pleistocene to Holocene Levant and their association with dietary habits. Journal of Archaeological Science: Reports, 22: 413-419, 2018.


M Edry, E Arie, H May, A Yasur-Landau. The Iron Age IIA Tombs of Area E, Tel Achziv: Between Local Traditions and the Consolidation of the Tyrian Polity. Israel Exploration Journal, 68(2): 150-181, 2018.

A Pokhojaev, W Habashi, H May, E Schulz-Kornas, N Shvalb, R Sarig. Examination of the Interproximal Wear Mechanism: Facet Morphology and Surface Texture Analysis. Journal of Dental Research, 97(13): 1145-1451, 2018.


Harney É, May H, Shalem D, Rohland N, Mallick S, Lazaridis I, Sarig R, Stewardson K, Nordenfelt S, Patterson N, Hershkovitz I, Reich D. Ancient DNA from Chalcolithic Israel reveals the role of population mixture in cultural transformation. Nat Commun., 9(1): 3336, 2018.

Sella-Tunis T, Pokhojaev A, Sarig R, O'Higgins P, May H. Human mandibular shape is associated with masticatory muscle force. Sci Rep., 8(1): 6042, 2018.

Hershkovitz I, Weber GW, Quam R, Duval M, Grün R, Kinsley L, Ayalon A, Bar-Matthews M, Valladas H, Mercier N, Arsuaga JL, Martinón-Torres M, Bermúdez de Castro JM, Fornai C, Martín-Francés L, Sarig R, May H, Krenn VA, Slon V, Rodríguez L, García R, Lorenzo C, Carretero JM, Frumkin A, Shahack-Gross R, Bar-Yosef Mayer DE, Cui Y, Wu X, Peled N, Groman-Yaroslavski I, Weissbrod L, Yeshurun R, Tsatskin A, Zaidner Y, Weinstein-Evron M. The earliest modern humans outside Africa. Science, 359(6734): 456-459, 2018.

Cohen H, Kugel C, May H, Medlej B, Stein D, Slon V, Brosh T, Hershkovitz I. The influence of impact direction and axial loading on the bone fracture pattern. Forensic Sci Int., 277: 197-206, 2017.

Tunis T, Sarig R, Cohen H, Medlej B, Peled N, May H. Sex estimation using computed tomography of the mandible. Int J Legal Med., 131(6): 1691-1700, 2017.

Cohen H, Kugel C, May H, Medlej B, Stein D, Slon V, Brosh T, Hershkovitz I. The effect of impact tool geometry and soft material covering on long bone fracture patterns in children. Int J Legal Med., 131(4): 1011-1021, 2017.

Abbas J, Slon V, May H, Peled N, Hershkovitz I, Hamoud K. Paraspinal muscles density: a marker for degenerative lumbar spinal stenosis? BMC Musculoskelet Disord, 17(1): 1-8, 2016.

Cohen H, Kugel C, May H, Medlej B, Stein D, Slon V, Hershkovitz I, Brosh T. The impact velocity and bone fracture pattern: Forensic perspective. Forensic Sci Int., 266: 54-62, 2016.


May H, Ruff C. Physical burden and lower limb bone structure at the origin of agriculture in the levant. AJPA., 161(1): 26-36, 2016.

Cohen H, Kugel C, May H, Medlej B, Stein D, Slon V, Hershkovitz I, Brosh T. The impact velocity and bone fracture pattern: Forensic perspective. Forensic Sci Int., 266:54-62, 2016.

Hay O, Dar G, Abbas J, Stein D, May H, Masharawi Y, Peled N, Hershkovitz I. The Lumbar Lordosis in Males and Females, Revisited. PLos One, 10(8): e.0133685, 2015.


Sarig R, Hershkovitz I, Shpack N, May H, Vardimon AD. Rate and pattern of interproximal dental attrition. Eur J Oral Sci., 123(4): 276-81, 2015.


Slon V, Stein D, Cohen H, Sella-Tunis T, May H, Hershkovitz I. Vertebral hemangiomas: their demographical characteristics, location along the spine and position within the vertebral body. Eur Spine J., 24(10): 2189-95, 2015.


H Cohen, V Slon, A Barash, H May, B Medlej, I Hershkovitz. Assyrian attitude towards captive enemies: A 2700-year-old paleo-forensic study. Int J Osteoarcheol., 25(3): 265-80, 2015.


Hershkovitz I, Donoghue HD, Minnikin DE, May H, Lee OY, Feldman M, Galili E, Spigelman M, Rothschild BM, Bar-Gal GK. Tuberculosis origin: The Neolithic scenario. Tuberculosis, 95: S122-6, 2015.


Hershkovitz I, Marder O, Ayalon A, Bar-Matthews M, Yasur G, Boaretto E, Caracuta V, Alex B, Frumkin A, Goder-Goldberger M, Gunz P, Holloway RL, Latimer B, Lavi R, Matthews A, Slon V, Mayer DB, Berna F, Bar-Oz G, Yeshurun R, May H, Hans MG, Weber GW, Barzilai O. Levantine cranium from Manot Cave (Israel) foreshadows the first European modern humans. Nature, 520(7546): 216-9, 2015.


May H, Rühli F. Is the Shape of the Proximal Femur a Risk Factor for Degenaritve Processes of the Hip and Hip Fracture? A New Approach Based On Geometric Morphometrics Methods. The FASEB Journal., 29 (1 Supplement): 702-3, 2015.


Hershkovitz I, Spigelman M, Sarig R, Lim DS, Lee IS, Oh CS, May H, Boaretto E, Kim YS, Lee SD, Peled N, Kim MJ, Toledano T, Bar-Gal GK, Shin DH. A possible case of cherubism in a 17th-century Korean mummy.

PLoS One, 9(8): e102441, 2014.


Sarig R, Hershkovitz I, Shvalb N, Sella-Tunis T, May H, Vardimon AD. Proximal attrition facets: morphometric, demographic, and aging characteristics. Eur J Oral Sci., 122(4): 271-8, 2014.


Slon V, Hershkovitz I, May H. The value of cadaver CT scans in gross anatomy laboratory. Anat Sci Educ., 7(1): 80-2, 2014.


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